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The PCM uses the fuel level sensor input in order to determine the amount of fuel in the fuel tank. The PCM disables the engine misfire diagnostic when the fuel level is too low and the EVAP system diagnostic when the fuel level is either too high or too low. The PCM also controls the fuel gauge based on the fuel level input.
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it really seems like if you want to use the 4th gen tank and have your fancy emissions crap connected and working, you need to use an ls1 cluster. i'm sure there's ways around everything if you don't wanna do that.